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Dr Phil Cox

Tel: 01904 321744    |    |   @drphilcox

Hull York Medical School, University of York, Heslington, York, YO10 5DD

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Cox PG, Hautier L (2015) Evolution of the Rodents: Advances in Phylogeny, Functional Morphology and Development. Cambridge: Cambridge University Press

Book Chapters

Hautier L, Cox PG (2015) Rodentia: a model order? In: Evolution of the Rodents: Advances in Phylogeny, Functional Morphology and Development (eds PG Cox, L Hautier). Cambridge: Cambridge University Press, pp. 1-18.

Hautier L, Cox PG, Lebrun R (2015) Grades and clades among rodents: the promise of geometric morphometrics. In: Evolution of the Rodents: Advances in Phylogeny, Functional Morphology and Development (eds PG Cox, L Hautier). Cambridge: Cambridge University Press, pp. 277-299.

Cox PG, Jeffery N (2015) The muscles of mastication and the function of the medial pterygoid. In: Evolution of the Rodents: Advances in Phylogeny, Functional Morphology and Development (eds PG Cox, L Hautier). Cambridge: Cambridge University Press, pp. 350-372.

Journal articles

PDF copies available on request: email

Cox PG (2017) The jaw is a second-class lever in Pedetes capensis (Rodentia: Pedetidae). PeerJ 5: e3741.

Davies TG, Rahman IA, Lautenschlager S, et al. (2017). Open data and digital morphology. Proceedings of the Royal Society B 284: 20170194.

Pataky TC, Koseki M, Cox PG (2016) Probabilistic biomechanical finite element simulations: whole-model classical hypothesis testing based on upcrossing geometry. PeerJ Computer Science 2: e96.

McIntosh AF, Cox PG (2016) The impact of digging on craniodental morphology and integration. Journal of Evolutionary Biology 29: 2383-2394.

McIntosh AF, Cox PG (2016) The impact of gape on the performance of the skull in chisel-tooth digging and scratch digging mole-rats (Rodentia: Bathyergidae). Royal Society Open Science 3: 160568. 

Gignac PM, Kley NJ, Clarke JA, Colbert MW, Morhardt AC, Cerio D, Cost IN, Cox PG, Daza JD, Early CM, Echols MS, Henkelman RM, Herdina AN, Holliday CM, Li Z, Mahlow K, Merchant S, Müller J, Orsborn C, Paluh DJ, Thies ML, Tsai HP, Witmer LM (2016) Diffusible iodine-based contrast-enhanced computed tomography (diceCT): an emerging tool for rapid, high-resolution, 3-D imaging of metazoan soft tissues. Journal of Anatomy 228: 889-909.

McIntosh AF, Cox PG, (2016) Functional implications of craniomandibular morphology in African mole-rats (Rodentia: Bathyergidae). Biological Journal of the Linnean Society 117: 447-462.

Cox PG, Baverstock H (2016). Masticatory muscle anatomy and feeding efficiency of the American beaver, Castor canadensis (Rodentia, Castoridae). Journal of Mammalian Evolution 23: 191-200.

Cox PGRinderknecht ABlanco RE (2015). Predicting bite force and cranial biomechanics in the largest fossil rodent using finite element analysis. Journal of Anatomy 226: 215-223.

Cox PG, Faulkes CG (2014). Digital dissection of the masticatory muscles of the naked mole-rat, Heterocephalus glaber (Mammalia, Rodentia). PeerJ 2: e448.

Cox PG, Kirkham J, Herrel A (2013). Masticatory biomechanics of the Laotian rock rat, Laonastes aenigmamus, and the function of the zygomaticomandibularis muscle. PeerJ 1: e160.


O'Hare LMS, Cox PG, Jeffery N, Singer ER (2013). Finite element analysis of stress in the equine proximal phalanx. Equine Veterinary Journal, 45: 273-277

Hautier L, Lebrun R, Cox PG (2012) Patterns of covariation in the masticatory apparatus of hystricognathous rodents: implications for evolution and diversification. Journal of Morphology 273: 1319-1337.

Cox PG, Rayfield EJ, Fagan MJ, Herrel A, Pataky TC, Jeffery N (2012). Functional evolution of the feeding system in rodents. PLoS ONE 7(4): e36299.

Cox PG, Fagan MJ, Rayfield EJ, Jeffery N (2011). Finite element modelling of squirrel, guinea pig and rat skulls: using geometric morphometrics to assess sensitivity. Journal of Anatomy 219: 696-709.

Cox PG, Jeffery N (2011). Reviewing the morphology of the jaw-closing musculature in squirrels, rats and guinea pigs with contrast-enhanced microCT. Anatomical Record Part A 294: 915-928. [cover article]

Jeffery NS, Stephenson R, Gallagher JA, Jarvis JE, Cox PG (2011). Micro-computed tomography with iodine staining resolves the arrangement of muscle fibres. Journal of Biomechanics 44: 189-192.

Jeffery N, Cox PG (2010). Do agility and skull architecture influence geometry of the mammalian vestibulo-ocular reflex? Journal of Anatomy 216: 496-509.

Cox PG, Jeffery N (2010). Semicircular canals and agility: the influence of size and shape data. Journal of Anatomy 216: 37-47.

Cox PG, Jeffery N (2008). Geometry of the semicircular canals and extraocular muscles in rodents, lagomorphs, felids and modern humans. Journal of Anatomy 213: 583-596.

Cox PG (2008). A quantitative analysis of the eutherian orbit: correlations with masticatory apparatus. Biological Reviews 83: 35-69. [cover article]

Cox PG, Jeffery N (2007). Morphology of the mammalian vestibulo-ocular reflex: the spatial arrangement of the human fetal semicircular canals and extraocular muscles. Journal of Morphology 268: 878-890.

Cox PG (2006). Character evolution in the orbital region of the Afrotheria. Journal of Zoology 269: 514-526


Hester Baverstock, Hull York Medical School

Ernesto Blanco, Instituto de Física, Montevideo


Michael Fagan, University of Hull

Chris Faulkes, Queen Mary University of London

Lionel Hautier, University of Montpellier

Anthony Herrel, Muséum National d'Histoire Naturelle, Paris

Nathan Jeffery, University of Liverpool

Michihiko Koseki, Shinshu University

Renaud Lebrun, University of Montpellier

Andrew McIntosh, Hull York Medical School

Todd Pataky, Shinshu University


Emily Rayfield, University of Bristol


Andrés Rinderknecht, Museo Nacional de Historia Natural, Montevideo

Ellen Singer, University of Liverpool

Artwork on Biological Reviews front cover by Jennifer May Hellier, King's College London